(quoted from post at 16:18:44 06/26/09) I worked on a farm that had 5 different 4000 SU's from 1971 to 1974 plus 3 or 4 4600 SU's. They ALL came new with 12x24 wheels AND 16.9x24 tires (it was just something that Ford did back then)! The 1973 had the fixed wheels (picture on right) while all the others had the adjustable wheels (picture on left). They are still running some of them 25 years later with that wheel and tire combination AND no problems!! My suspision is that the fixed center wheels are cheaper and would be found on a tractor that didn't require as much adjustment flexibility. The fixed center wheels will give you a tread width of 60 inches dished in and a tread width of 72 inches dished out. the adjustable center wheels can be adjusted in 4 inch steps from 56 inches to 78 inches, (if memory serves). HTH. Oh yeah, one other thing! They NEVER had a wheel failure with either type of wheel!
Here's a picture of a 16.9x24 tire on a 12x24 rim
Edit addendum, these tires were ALL R-1's AND these tractors WORKED hard! (Plowing, discing, pulling trailers with 5 ton loads, orchard spraying, etc.)
Another thought -- If you want the fixed center wheel, you will probably find it quicker on a light industrial (like a 515 or similar). I would think that you are more likely to find the adjustable wheel on an ag tractor!